Журнал Сибирского федерального университета
Journal of Siberian Federal University
Биология
Biology
Редакционный совет:
академик РАН Е.А. Ваганов
академик РАН И.И. Гительзон
академик РАН А.Г. Дегерменджи
академик РАН В.Ф. Шабанов
чл.-корр. РАН, д-р физ.-мат. наук
В.Л. Миронов
чл.-корр. РАН, д-р техн. наук
Г.Л. Пашков
чл.-корр. РАН, д-р физ.-мат. наук
В.В. Шайдуров
чл.-корр. РАН, д-р физ.-мат. наук
В.В. Зуев
Editorial Advisory Board
Chairman:
Eugene A. Vaganov
Members:
Josef J. Gitelzon
Vasily F. Shabanov
Andrey G. Degermendzhy
Valery L. Mironov
Gennady L. Pashkov
Vladimir V. Shaidurov
Vladimir V. Zuev
Editorial Board:
Editor-in-Chief:
Mikhail I. Gladyshev
Founding Editor:
Vladimir I. Kolmakov
Managing Editor:
Olga F. Alexandrova
Executive Editor for Biology:
Elena S. Kravchuk
CONTENTS / СОДЕРЖАНИЕ
Vera Yu. Kovaleva,
Vadim M. Efimov and Yuri N. Litvinov
Directional Asymmetry of Morphological Traits During Postnatal
Ontogeny in Root Vole Microtus oeconomus Pall. (Rodentia,
Cricetidae)
– 115 –
Е.П. Турпаева, А.К. Райский
l%!“*,е
C=3*,
!%д=
Colossendeis
(Colossendeidae,
Pycnogonida) м%! r.дделл= , C!,ле›=?,. =*"=2%!,L
– 130 –
А.А. Савченко, К.П. Базарин
q%“2% …,е
=*2,"…%“2, m`d- , m`dt-ƒ=",“,м/.
дег,д!%ге…=ƒ " …еL2!%-,ль…/. г!=…3л%ц,2=. 3 “C%!2“ме…%"
" д,…=м,*е 2!е…,!%"%ч…%г% ц,*л=
– 151 –
Â.Ñ. Áîíäàðü, À.Ï. Ïóçûðü,
À.Â. Ãîðåâà, À.Å. Буров
j%…“2!3,!%"=…,е *%мC%ƒ,ц,%……/. м=2е!,=л%" …= %“…%"е
…=…%=лм=ƒ%" , !=ƒ!3ш=ем/. K,%C%л,ме!%"
– 163 –
Ï.À. Øåñòåðíÿ, À.Ñ. Ñåðãååâà,
С.Ю. Никулина, В.А.Шульман
k%*3“ 9!21.3 $ ге…е2,че“*,L C!ед,*2%! 2 ›е“2, *%!%…=!…%г%
=2е!%“*ле!%ƒ= 3 м3›ч,…
– 175 –
Редактор И.А. Вейсиг Корректор Е.Г. Иванова
Компьютерная верстка Е.В. Гревцовой
Подписано в печать 28.06.2013 г. Формат 84x108/16. Усл. печ. л. 8,7.
Уч.-изд. л. 8,2. Бумага тип. Печать офсетная. Тираж 1000 экз. Заказ 3134.
Отпечатано в ПЦ БИК. 660041 Красноярск, пр. Свободный, 82а.
2013 6 (2)
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Editorial board for Biology:
Elena Kravchuk – Series Editor, Institute of Biophysics,
Russian Academy of Sciences, Siberian Branch,
Russia
Nadezhda Sushchik – Institute of Biophysics, Russian
Academy of Sciences, Siberian Branch, Russia
Sergey Bartsev – Institute of Biophysics, Russian
Academy of Sciences, Siberian Branch, Russia
Alexander Bolsunovsky – Institute of Biophysics,
Russian Academy of Sciences, Siberian Branch,
Russia
Marc d’Alarcao – Tufts University, USA
Nicolai Gaevsky – Siberian Federal University, Russia
Woodland J. Hastings – Harvard University, USA
Malcolm Hughes – the University of Arizona, USA
Takayoshi Koike – Hokkaido University, Japan
Valentina Kratasyuk – Siberian Federal University;
Institute of Biophysics, Russian Academy of
Sciences, Siberian Branch, Russia
Elena Muratova – Institute of Forest, Russian Academy
of Sciences, Siberian Branch, Russia
Akira Osawa – Kyoto University, Japan
Frank B. Salisbury – NASA, USA
Ernst-Detlef Shulze – Max Planck Institute of
Biogeochemistry, Germany
Tatyana Volova – Siberian Federal University; Institute
of Biophysics, Russian Academy of Sciences,
Siberian Branch, Russia
Eugene Vysotsky – Institute of Biophysics, Russian
Academy of Sciences, Siberian Branch, Russia
Egor Zadereev – Institute of Biophysics, Russian
Academy of Sciences, Siberian Branch, Russia
Liliana Zalizniak – RMIT University, Australia
Ramesh Gulati – NIOO/Centre of Limnology, The
Netherlands
Mikhail Karpinsky – Russian Federal Research Institute
of Fisheries and Oceanography, Russia
Ã.Ò. Îìóðîâà, Â.Â. Áàðèíîâ,
Î.Â. Êàðäàø, Â.Ñ. Мыглан
r“2=…%"ле…,е "!еме…, “2!%,2ель“2"= (Cе!е“2!%L*,)
m=д/м“*%г%
=“Cе*2
г%!%д*=:
– 185 –
Е.Г. Крылова
r“2%Lч,"%“2ь
C!ед“2=",2елеL !%д=
– 196 –
Т.Т. Ефремова, С.П. Ефремов
o!,!%д= *,“л%2…/. “"%L“2" C%д“2,л*, K%л%2…/.
Kе!еƒ… *%"
– 205 –
Bidens
(Asteraceae) * деL“2",ю “3ль-=2= ц,…*= …=
…=ч=ль…/. .2=C=. %…2%ге…еƒ=
де…д!%.!%…%л%г,че“*,L
Свидетельство о регистрации СМИ
ПИ № ФС77-28-725 от 29.06.2007 г.
Серия включена в «Перечень ведущих рецензируемых
научных журналов и изданий, в которых
должны быть опубликованы основные
научные результаты диссертации на соискание
ученой степени доктора и кандидата наук» (редакция
2010 г.)
Стр.2
Journal of Siberian Federal University. Biology 2 (2013 6) 115-129
~ ~ ~
УДК 574.3, 591.3
Directional Asymmetry of Morphological
Traits During Postnatal Ontogeny
in Root Vole Microtus oeconomus Pall.
(Rodentia, Cricetidae)
Vadim M. Efi mova,b,c
a
Vera Yu. Kovalevaa
and Yuri N. Litvinova
*,
Institute of Systematics and Ecology of Animals SB RAS,
11 Frunze Str., Novosibirsk, Russia 630091
bInstitute of Cytology and Genetics SB RAS,
10 Pr. Lavrentyeva, Novosibirsk, Russia 630090
c
Tomsk State University,
36 Pr. Lenina, Tomsk, Russia 634050
Received 05.11.2012, received in revised form 22.11.2012, accepted 19.05.2013
We analyzed Microtus oeconomus material collected for 25 years (1982–1992, 1994–2005,
2009, 2010). Directional asymmetry (DA) was demonstrated for teeth morphotypes, adrenal and
kidney weights, and for length of the hind paw as well. DA may be formed both at early and
later development stages and is not dependent on environmental factors and population density
dynamics. Although each trait has its own dynamics DA in ontogeny, usually DA increases in
the period of sexual maturation. Uncoordinated development of the sides in this period seems
to associate with an increase in intensity of developmental processes. Our study argues that
sexual dimorphism of continuous traits appears at the later developmental stages. This fi nding
is consistent with Geodakian’s evolutionary theory of sex. Evolutionary signifi cance of the DA
is to increase both short-term individual fi tness and long-term population adaptive capacity.
Developmental approach to the study of morphological and functional asymmetries in animals
opens new possibilities for the study of evolutionary processes.
Keywords: Microtus oeconomus, directional asymmetry, teeth morphotypes, adrenal and kidney
weights, paw lengths, environmental conditions, population density dynamics.
© Siberian Federal University. All rights reserved
* Corresponding author E-mail address: vkova@ngs.ru
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